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In addition, microbial metabolites critically shape IFN-γ programs: short-chain fatty acids (SCFAs) modulate IL-12 production by dendritic cells and promote CD8 + T-cell effector differentiation (94, 95), and indole derivatives regulate mucosal immune homeostasis through AHR signaling, thereby influencing IFN-γ expression in Th1 and NK cells (96, 97)